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Transcripción

1 Facultad de Ciencias! " # $%&'()'!*' +&*+'(,- Tesis presentada por Álvaro Sanz Sáez de Jáuregui para optar al Grado de Doctor por la Universidad de Navarra

2 Servicio de Publicaciones de la Universidad de Navarra ISBN

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4 F a c u l t a d d e C i e n c i a s! " # $%&'()'!*' +&*+'(,- Memoria presentada por D. Álvaro Sanz Sáez de Jáuregui para aspirar al grado de Doctor en Ciencias Biológicas por la Universidad de Navarra El presente trabajo ha sido realizado bajo nuestra dirección en el Departamento de Biología Vegetal, Sección Biología Vegetal y autorizamos su presentación ante el Tribunal que lo ha de juzgar. Pamplona, 1 de Septiembre de 2011 Dr. Juan José Irigoyen Iparrea Dr. Gorka Erice Soreasu

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44 WEEK 2 WEEK 4 TREATMENTS Total Dry Weight Nodule Dry Weight Total Leaf Area Root/shoot ratio SLA Total Dry Weight Nodule Dry Weight Total Leaf Area Root/shoot ratio SLA 0A 0.32 ± 0.11 b 7.6 ± 1.5 a 17.7 ± 2.2 c 0.44 ± ± 12.9 bc 1.04 ± 0.21 d 31,1 ± 8,5 a ± 9.8 c 0.46 ± ± 18.8 a 0E 0.14 ± 0.02 c 4.6 ± 1.2 b 11.7 ± 2.1 c 0.41 ± ± 3 d 1.37 ± 0.40 d 34,1 ± 2,5 a 69.5 ± 7.2 c 0.31 ± ± 12.5 c 10A 0.44 ± 0.03 ab 1.1 ± 0.2 c 56.2 ± 3.6 a 0.31 ± ± 19.4 a 2.83 ± 0.29 c 0,9 ± 0,1 b ± 22 ab 0.36 ± ± 7.9 a 10E 0.62 ± 0.04 a 0.8 ± 0.1 c 56.6 ± 6 a 0.39 ± ± 7.1 c 4.08 ± 0.15 ab 11,3 ± 0,7 b 293 ± 9.8 ab 0.36 ± ± 9.1 b 15A 0.47 ± 0.10 ab 1.3 ± 0.4 c 39.1 ± 8.4 b 0.26 ± ± 7.8 b 3.28 ± 0.16 bc 0,01 ± 0,0014 b ± 19.4 a 0.38 ± ± 5.3 a 15E 0.48 ± 0.07 ab 1 ± 0.04 c 49.3 ± 7.4 ab 0.35 ± ± 16.5 bc 4.56 ± 0.82 a 0,2 ± 0,0025 b ± 27.1 b 0.35 ± ± 18.9 b CO 2 ns ns ns ns *** * ns ns ns *** NH 4NO 3 ** *** *** ns *** *** *** *** ns ** CO 2*NH 4NO 3 ns ns ns ns ** ns ns ns ns ns $% * + ; $'! ' [ " 0225 *!*# [ & )/ 0 ) 1 $2! 2 3 4& ; + $ &! ; + $ &! $ &! 9+ $,'! & + 00 $A< & D2 $A< 0& 8 + Q *@ [ 0 ; ; ' + 8 $')8'&! =+ ')8';>\! 3B \\ B! \\\ 2B A+ ; ')8'!,7 ; 4 ; + $- ] 223&,7 A+ ; ')8'!,7 ;! + ; 1

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46 WEEK 2 WEEK 4 TREATMENTS Vc max A growth Ci Vc max A growth Ci 0A ± ± 6.5 b ± 6.2 c 94.6 ± 2.9 b 20 ± 6.5 b ± 18.5 b 0E 98.1 ± ± 9.6 a ± 6.9 ab 82.2 ± 6.7 c 24.6 ± 9.6 ab ± 26.3 a 10A ± ± 1.2 b ± 27.6 c ± 5 a 22.6 ± 1.2 ab ± 16.2 b 10E ± ± 3.6 ab ± 17.1 a 97.6 ± 6 ab 25.1 ± 3.6 ab 411 ± 39 a 15A ± 7, ± 8.8 b ± 12.0 c ± 15.4 a 21 ± 8.8 b ± 19.7 b 15E ± 8, ± 1.8 ab ± 13.9 b ± 5.9 a 29.1 ± 1.8 a 412 ± 10 a CO 2 ns *** *** * ** *** NH 4NO 3 ns * * * ns ns CO 2*NH 4NO 3 ns ns ns ns ns ns $ * + ; $'! ' [ " 0225 *! *# [ & )/ 0 ) 1 $2! 2 3 4& " $8 "! 5 &! ++ ;+ $' ;+! 5 & + $! 5 &! + 00 $A< & D2 $A< 0& 8 + Q *@ [ 0 ; ; ' + 8 $')8'&! =+ ')8';>\! 3B \\ B! \\\ 2B A+ ; ')8'!,7 ; 4 ; + $- ] 223&,7 A+ ; ')8'!,7 ;! + ; ' + ;<! " # " $8 " &; + )/ 0 ) 1 $= & )#+! + + ;<! # 8 " )/ 0 ) 1 $,&' " & ( "( $$" & + + )/ 0 ) 1 $&' " & ( (!( $$"!& *# + + ;< + $@ "%#*" + $$$%.!! + $$$% #& $= & ' + ;< + ; + )/ 0 ) 1 $&' " & ( ()( $$(& +! + + ;<)/ 0 ) 1 + $= & 11

47 ' + ;<! + =- $= 1& ; + # ;+ + ; ;+ 3 4 )/ 0 ) 1 )#+! ;+ ; ;+ 2 4 )/ 0 ) 1! =- $,&' " & ( " $$$%& ' + + ;<! =- ; )/ 0 ) 1 ' + ;<! ++ ) $-)*& $= 1& ; + ;+ 24 )/ 0 ) 1 ; # $, &' " & ( *%%"! $$& ' + + ;<! ;+ 24 )/ 0 ) 1 ; # + + -)* $= 1& #! ;+ 3 4 )/ 0 ) 1 ; # -)* $,&' " & ( ")#% $$!& ' + ;<! + $=& ; + ;+ 2 4 )/ 0 ) 1 $= 1& - ; ; )/ 0 ) 1 +; ; = + ;+ 2 4 )/ 0 ) 1 $,&' " & ( %" $$$#& TREATMENTS WEEK 2 WEEK 4 TSP PNUE TSS Starch TSP PNUE TSS Strach 0A 3.2 ± 0.2 ab 14.3 ± 0.6 b 0.9 ± 0.10 bc 2.8 ± 0.10 b 2.3 ± ± 0.3 bc 1.3 ± 0.02 b 1.4 ± 0.10 b 0E 1.8 ± 0.2 c 24.3 ± 2.3 a 1.1 ± 0.08 ab 5.5 ± 0.30 a 2.0 ± ± 1.0 ab 1.8 ± 0.09 a 2.4 ± 0.60 a 10A 1.8 ± 0.2 c 15.3 ± 2.8 b 0.6 ± 0.04 e 1.8 ± 0.07 c 2.8 ± ± 1.4 a 0.8 ± 0.05 d 0.6 ± 0.20 c 10E 2.3 ± 0.4 bc 15.0 ± 1.2 b 1.3 ± 0.10 a 2.3 ± 0.30 b 2.5 ± ± 0.4 ab 1.7 ± 0.07 a 2.7 ± 0.10 a 15A 2.0 ± 0.3 c 13.5 ± 1.5 b 0.7 ± 0.03 de 1.4 ± 0.06 c 2.5 ± ± 0.6 ab 1.0 ± 0.05 cd 0.6 ± 0.20 c 15E 3.6 ± 0.4 a 14.8 ± 1.5 b 0.9 ± 0.03 cd 1.8 ± 0.03 c 1.9 ± ± 0.3 c 1.1 ± 0.10 bc 0.2 ± 0.06 c CO 2 ns * *** *** ns ns *** *** NH 4 NO 3 ns * * *** ns * *** *** CO 2 *NH 4 NO 3 *** * *** ** ns * *** *** $) * + ; $'! ' [ " *! *# [ & )/ 0 ) 1 $2! 2 3 4& $=-! &! ++ ) $-)*! 5 ) & $=! &! + $ & # + 00 $A< & D2 $A< 0& 8 + Q *@ [ 0 ; ; ' + 8 $')8'&! =+ ')8' ;>\! 3B \\ B! \\\ 2B A+ ; ')8'!,7 ; 4 ; + $- ] 223&,7 10

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