Libro de resúmenes Abstract book 1
Editado por: Fidel Torcida Fernández-Baldor y Pedro Huerta De esta edición: Colectivo Arqueológico y Paleontológico de Salas, C.A.S. 2013 Publica: Colectivo Arqueológico y Paleontológico de Salas, C.A.S. Plaza Jesús Aparicio 9, 1º 09600 Salas de los Infantes (Burgos) E-mail: secretaria@colectivosalas.com Depósito Legal: BU 252-2013 ISBN - 10: 84-695-8567-3 ISBN - 13: 978-84-695-8567-2 2
LIBRO DE RESÚMENES ABSTRACTS BOOK VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno VI International Symposium about Dinosaurs Palaeontology and their Environment Salas de los Infantes, 5-7 de septiembre de 2013 3
COMITÉ CIENTÍFICO Ainara Badiola Universidad del País Vasco/EHU, España. J. Luis Barco Paleoymas. Zaragoza, España. Ángela Delgado Buscalioni Universidad Autónoma de Madrid, España. Juan Canale Museo Ernesto Bachmann, El Chocón, Argentina. J. L. Carballido Museo Egidio Feruglio, Trelew, Argentina. Julio Company Universidad Politécnica de Valencia, España. Rodolfo Coria Universidad Nacional de Río Negro, Argentina. Penélope Cruzado CONICET-Inst. de Investigac. en Paleobiología y Geología, Argentina. Gloria Cuenca Universidad de Zaragoza, España. J. Bienvenido Diez Ferrer Universidad de Vigo, España. James Farlow Indiana Purdue University, USA. Pascal Godefroit Royal Belgian Institute of Natural Sciences, Bélgica. Joaquín Moratalla Instituto Geológico y Minero de España, España. Adán Pérez García Universidad Complutense de Madrid, España. Félix Pérez Lorente Universidad de la Rioja, España. Rafael Royo-Torres Fundc. Conjunto Paleontológico de Teruel-Dinópolis, España. Luis Miguel Sender Universidad de Zaragoza, España. José Ignacio Ruiz-Omeñaca Museo del Jurásico de Asturias, España. COMITÉ ORGANIZADOR Coordinadores: Fidel Torcida Fernández-Baldor, C.A.S. y Museo de Dinosaurios (Salas de los Infantes, Burgos). Xabier Pereda Suberbiola, Universidad del País Vasco/EHU, Bilbao. José Ignacio Canudo Sanagustín, Universidad de Zaragoza, Grupo Aragosaurus. Pedro Huerta Hurtado, C.A.S. y Universidad de Salamanca. Miembros: Víctor Urién Montero, C.A.S. y Museo Dinosaurios, Salas de los Infantes. Gustavo Pérez Martínez, C.A.S. y Museo Dinosaurios, Salas de los Infantes. Diego Montero Huerta, C.A.S. y Museo Dinosaurios, Salas de los Infantes. Alberto Bengoechea Molinero, C.A.S. y Museo Dinosaurios, Salas de los Infantes. Rubén Contreras Izquierdo, C.A.S. y Museo Dinosaurios, Salas de los Infantes. Silvia Mielgo Gallego. Asier Pascual Marquínez, C.A.S. y Museo Dinosaurios, Salas de los Infantes. Damián González Oyagar, C.A.S. Manuel García Ávila, Universidad de Vigo. Colaboradores: Sergio Urién Montero, Museo Dinosaurios, Salas de los Infantes. Luis Ángel Izquierdo Montero, Museo Dinosaurios, Salas de los Infantes. Cecilia Heras. Natalie Bardet, Muséum National d Histoire Naturelle de Paris (Francia). Antonio Rodriguez Verísimo, Museo de Dinosaurios de Salas de los Infantes. Evangelina Alonso Alzaga, Keith James. José Ignacio Díaz Martínez, Universidad de La Rioja. 4
Contenido Programa/Schedule 9 Presentación/Presentation 13 Conferencias/Keynotes A cross-perspective into the Early Cretaceous dinosaur faunas from Europe and Asia: new data on the dinosaurs from the Savannakhet Bassin (Laos) and from the lignitic bonebed of Angeac-Charente (France). Allain, R., Rozada. L., Vullo, R. 17 Ten years digging up dinosaurs in Patagonia: the Ibero-Patagonian connection in the Mesozoic. Canudo, J. I. Salgado, L. Carballido, J. Garrido, A. 19 Sauropod herbivory and the Mesozoic flora. Gee, C. T. 21 Overview on dinosaur tracksites in the Swiss and French Jura Mountains. Marty, D., Meyer, C.A., Paratte, G. Cattin, M. 23 Towards the last European ecosystems with dinosaurs: the Upper Cretaceous biota of Lo Hueco (Fuentes, Cuenca, Spain). Ortega, F. 27 Rebbachisauridae: the European exception. Torcida Fernández-Baldor, F., Canudo, J.I. 31 The dinosaurs of the Pyrenees: distribution and palaeoenvironments. Vila, B. 33 The new giant pterosaurs: novel anatomies and habits in the largest flying animals. Witton, M.P. 37 Comunicaciones/Communications. An eusuchian skeleton from the lower Maastrichtian palustrine deposits of Fumanya (south-eastern Pyrenees, Iberian Peninsula). Blanco, A., Puértolas-Pascual, E., Vila, B., Marmi, J. 43 Europasaurus holgeri the dwarfed dinosaur from the Late Jurassic of Germany. Bolecsek, C., Wings, O. 46 The discovery and loss of the colossal Brontosaurus giganteus from the fossil fields of Wyoming (USA) and the events that led to the discovery of Diplodocus carnegii: the first mounted dinosaur on the Iberian Peninsula. Breithaupt, Brent H. 48 Tracking ornithopod tracks in the Jurassic-Cretaceous interval of the Iberian Range (Spain). Castanera, D., Razzolini, N.L., Vila, B., Pascual, C., Barco, J.L., Canudo, J.I. 51 A diminutive hadrosaur from the Late Maastrichtian of the south-central Pyrenees (Spain). Company, J., Cruzado-Caballero, P. and Canudo, J.I. 53 Utilidad de la tomografía computerizada (CT) en la preparación paleontológica de un fósil de vertebrado del Cretácico Inferior (Mesozoico). Contreras-Izquierdo, R., Torcida Fernández-Baldor, F., Castilla, M., Rodríguez, L. y Santos, E. 56 5
Mammals from the time of the dinosaurs, the symmetrodontans from the Early Cretaceous of Spain. Cuenca-Bescós, G., Canudo, J.I., Gasca, J.M., Moreno-Azanza, M., Cifelli, R.L. 58 Un nuevo yacimiento con icnitas de dinosaurio en el Grupo Urbión (Cuenca de Cameros, España). Díaz-Martínez, I., García-Ortiz, E. y Pérez-Lorente, F. 60 Phylogenetic relationships of the titanosaurian dinosaurs from the Late Cretaceous of the Ibero-Armorican Island. Díez Díaz, V., Garcia, G., Pereda Suberbiola, X., Valentin, X., and Sanz, J.L. 63 Parasitism in dinosaur clutches? Fernández, M.S. and García, R.A. 65 XL Hadrosaurids of Tremp basin: the Costa de Les Solanes site (Late Maastrichtian, northeast of the Iberian Peninsula). Fondevilla, V., Razzolini, N. L., Gaete, R., Pellicer Mir, X., Galobart, A. 67 Palaeoecological reconstruction of the late Hauterivian-early Barremian fossil site of Horcajuelos (Salas de los Infantes, Burgos province, Spain). García Ávila, M., Sender, L.M., Villanueva-Amadoz, U., Torcida Fernández-Baldor, F., Diez, J.B. 69 New fossil remains of the ornithopod dinosaur Delapparentia turolensis (Early Cretaceous, Spain) and its phylogenetic position. Gasca, J.M., Moreno-Azanza, M., Ruiz-Omeñaca, J.I., Canudo, J.I. 71 Dinosaur fossil sites from Ladruñán anticline (Mirambel Formation, Early Cretaceous, Teruel, Spain): tracks and bonebeds. Gasca, J.M., Moreno-Azanza, M., Canudo, J.I, Díaz-Martínez, I., Castanera, D., Ramón del Río, D. 74 A reappraisal of the spinosaurid baryonychines from the Arcillas de Morella Formation (lower Aptian) of Morella, Spain. Gasulla, J.M., Escaso, F., Ortega, F. and Sanz, J.L. 76 Palaeontology, Pangaea and Plate Tectonics Data before Models. James, H. K. 78 Extraction proposal and enhancement of a cretaceous Bennettital locality in Rabanera del Pinar (Burgos, Spain). Juncal Rosales, M.A., Sender Palomar, L.M., García Ávila, M., Torcida Baldor, F., Lanaja del Busto, J.M., Diez Ferrer, J.B. 79 Rediscovery of a lost portion of the holotype of Suchosaurus girardi (Sauvage, 1897-98), now related to the spinosaurid theropod Baryonyx Malafaia, E.; Ortega, F.; Escaso, F.; Mocho, P. 82 The Taller de Empleo de Restauración Paleontológica de Lo Hueco (Cuenca, Spain). Marcos, F.; Jiménez, M.V.; Valverde, R.; Abós, E.; Antón, E.; Calvo, C.; Estevan, R.; Igualada, C.; Llandres, M.; Toledano, M.I.; Parra, A.; Peña, C. 85 Megafossil plant assemblages from the early Maastrichtian of northeastern Iberia challenge the paradigm of the rise to dominance of angiosperms at local scales Marmi, J., Villalba-Breva, S., Gomez, B., Martín-Closas, C., and Daviero-Gomez, V. 86 Spinophorosaurus (Sauropoda), a new look inside eusauropod evolution Mocho, P., Ortega, F., Aberasturi, A., Escaso, F. 89 6
New approach to Lourinhasaurus alenquerensis (Macronaria, Camarasauromorpha) from the Portuguese Upper Jurassic. Mocho, P; Royo-Torres, R., & Ortega, F. 91 Ootaxa assemblage from El Horcajo site (upper Berriasian, Cameros Basin) in Trevijano (La Rioja, Spain). Moreno-Azanza, M., Gasca, J.M., Díaz-Martínez, I. Fernández, A., Canudo, J.I, Pérez-Lorente, F. 93 A new basal eusuchian crocodile from the Late Cretaceous of Lo Hueco (Cuenca, Spain). Narváez, I., Ortega, F., Brochu, C. & Escaso, F. 96 A new sauropod braincase from Azenak (Agadez, Niger) Páramo, A., Escaso, F., Aberasturi, A., Ortega, F. 98 Postcranial elements of Maledictosuchus riclaensis (Thalattosuchia) from Callovian of Spain. Parrilla-Bel, J. Canudo, J. I. 99 First dinosaur and turtle remains from the latest Cretaceous shallow marine deposits of Albaina (Laño Quarry, Condado de Treviño, Iberian Peninsula). Pereda Suberbiola, X. Corral, J.C. Murelaga, X. Martin, G. Larrañaga, J. Bardet, N. Pérez-García A., Berreteaga, A. 102 Un nuevo miembro basal de Eucryptodira (Chelonii) en el Jurásico Superior de Portugal Pérez-García, A., Ortega, F. 104 Nuevos datos sobre Pelobatochelys, Enaliochelys y otros miembros basales de Eucryptodira del registro del Jurásico Superior británico Pérez-García, A. 107 Una nueva hipótesis sobre las relaciones filogenéticas y la distribución paleobiogeográfica de los miembros de Paracryptodira, un abundante y diverso clado de tortugas del Mesozoico y Paleógeno norteamericano y europeo. Pérez-García, A. 110 La crisis finicretácica supuso la extinción de los linajes de tortugas primitivas (stem Testudines) del Mesozoico de Europa?. Pérez-García, A. 112 Dermal skeleton of a lower Albian goniopholidid (Crocodylomorpha, Neosuchia) from Andorra (Teruel, Spain): biomechanical implications. Puértolas-Pascual, E., Canudo, J.I. 115 Unusual dinosaur tracksite on Albian (Escucha Formation) soft sediments in Oliete. (Teruel, Spain). Royo-Torres, R., González, A., Mampel, L., Gascó, F., Ayala, D., Cobos, A., Espílez, E., Verdú, J. & Alcalá, L. 118 Sauropod tracks from the Early Cretaceous of Parede beach (Cascais, Portugal). Santos, V.F., Castanera, D., Barroso-Barcenilla, F., Callapez, P.M., Cupeto, C.A. and Rodrigues, N.P.C. 120 Laying under stressful conditions: temporal occurrence of pathologic sauropod eggshells in the Late Cretaceous of northeastern Iberian Peninsula Sellés, A. G. and Vila, B. 123 New contributions to the palaeobotany of Arroyo de la Vega fossil site (Early Cretaceous) at Rabanera del Pinar village (Burgos, Spain). Sender, L.M., Villanueva-Amadoz, U., García Ávila, M., Diez, J.B., Torcida Fernández-Baldor, F., Huerta, 7
P., Ferrer, J. 125 Descripción de icnitas de dinosaurios cuadrúpedos del tránsito Jurásico-Cretácico (Burgos, España). Torcida Fernández-Baldor, F., Díaz-Martínez, I, Contreras, R., Huerta, P., Montero D., Urién, V. 127 Description of a dorsal vertebra from the sauropod from El Oterillo II (Burgos, Spain). Torcida Fernández-Baldor, F., Canudo, J.I., Contreras, R., Huerta, P., Montero, D., Urién, V. 130 Reconstructing the environment and foraging behaviour of the latest titanosaur herds (Fumanya tracksite, NE Spain). Vila, B., Marmi, J., Villalba-Breva, S., Martín-Closas, C. 132 8
Programa/Schedule 9
Horario Jueves día 5 8:30 Recepción de asistentes y entrega de documentación/reception of participants 9:15 Presentación de las Jornadas 9:30 Conferencia Plenaria/Keynote lecture Jose Ignacio Canudo. Ten years digging up dinosaurs in Patagonia: the Ibero-Patagonian connection in the Mesozoic. 10:15 Sesión de comunicaciones/oral session Programa/Schedule 10:15-10:30. Narváez, I., Ortega, F., Brochu, C. y Escaso, F. A new basal eusuchian crocodile from the Late Cretaceous of Lo Hueco (Cuenca, Spain) 10:30-10:45. Pérez-García, A., Ortega, F. Un nuevo miembro basal de Eucryptodira (Chelonii) en el Jurásico Superior de Portugal 10:45-11:00. Pereda Suberbiola, X., Corral, J.C., Murelaga, X., Martin, G., Larrañaga, J., Bardet, N. Pérez-García A., Berreteaga, A. First dinosaur and turtle remains from the latest Cretaceous shallow marine deposits of Albaina (Laño Quarry, Condado de Treviño, Iberian Peninsula) 11:00 Pausa café and posters/cofee break and posters 11:30-11:45. Puértolas-Pascual, E., Canudo, J.I. Dermal skeleton of a lower Albian goniopholidid (Crocodylomorpha, Neosuchia) from Andorra (Teruel, Spain): biomechanical implications. 11:45-12:00. Parrilla-Bel, J.1 Canudo, J. I. Postcranial elements of Maledictosuchus riclaensis (Thalattosuchia) from Callovian of Spain. 12:00-12:15. Marcos, F.; Jiménez, M.V.; Valverde, R.; Abós, E.; Antón, E.; Calvo, C.; Estevan, R.; Igualada, C.; Llandres, M.; Toledano, M.I.; Parra, A.; Peña, C. The Taller de Empleo de Restauración Paleontológica de Lo Hueco (Cuenca, Spain) 12:15-12:30. Breithaupt, B. H. The discovery and loss of the colossal Brontosaurus giganteus from the fossil fields of Wyoming (USA) and the events that led to the discovery of Diplodocus carnegii: the first mounted dinosaur on the Iberian Peninsula. 12:30 Conferencia Plenaria/Keynote lecture Daniel Marty. Overview on dinosaur tracksites in the Swiss and French Jura Mountains 13:15 Actos Protocolarios/Protocolary acts. 14:00 Comida/Lunch time 16:00 Conferencia Plenaria/Keynote lecture Mark Witton. The new giant pterosaurs: novel anatomies and habits in the largest flying animals. 16:45 Sesión de comunicaciones/oral session 16:45-17:00. Pérez-García, A. Una nueva hipótesis sobre las relaciones filogenéticas y la distribución paleobiogeográfica de los miembros de Paracryptodira, un abundante y diverso clado de tortugas del Mesozoico y Paleógeno norteamericano y europeo. 17:00-17:15 Castanera, D., Razzolini, N.L., Vila, B., Pascual, C., Barco, J.L., Canudo, J.I. Tracking ornithopod tracks in the Jurassic-Cretaceous interval of the Iberian Range (Spain) 17:15-17:30. Díaz-Martínez, I., García-Ortiz, E. y Pérez-Lorente, F. Un nuevo yacimiento con icnitas de dinosaurio en el Grupo Urbión (Cuenca de Cameros, España). 17:30 Pausa café/cofee break and posters 18:00-18:15. Gasca, J.M., Moreno-Azanza, M., Canudo, J.I, Díaz-Martínez, I., Castanera, D., Ramón del Río, D. Dinosaur fossil sites from Ladruñán anticline (Mirambel Formation, Early Cretaceous, Teruel, Spain): tracks and bonebeds 18:15-18:30. Torcida Fernández-Baldor, F., Díaz-Martínez, I, Contreras, R., Huerta, P., Montero D., Urién, V. Descripción de icnitas de dinosaurios cuadrúpedos del tránsito Jurásico-Cretácico (Burgos, España). 18:30-18:45. Moreno-Azanza, M., Gasca, J.M., Díaz-Martínez, I., Fernández, A., Canudo, J.I, Pérez-Lorente, F. Ootaxa assemblage from El Horcajo site (upper Berriasian, Cameros Basin) in Trevijano (La Rioja, Spain). 18:45 Conferencia plenaria/keynote lecture Francisco Ortega. Towards the last European ecosystems with dinosaurs: the Upper Cretaceous biota of Lo Hueco (Fuentes, Cuenca, Spain). 19:30 Final de la sesión 20:30 Actividades culturales/cultural activities Conferencia de M.A. Sabadell, Editor Científico de Muy Interesante, El mundo perdido del Mesozoico: ciencia, sociedad y medios de comunicación. En el Teatro Auditorio Gran Casino. 10
Horario Viernes día 6 9:15 Conferencia Plenaria/Keynote lecture Bernat Vila. The dinosaurs of the Pyrenees: distribution and palaeoenvironments. 10:00 Sesión de comunicaciones/oral session 10:00-10:15. Fondevilla, V., Razzolini, N. L., Gaete, R., Pellicer Mir, X., Galobart, A XL Hadrosaurids of Tremp basin: the Costa de Les Solanes site (Late Maastrichtian, northeast of the Iberian Peninsula). 10:15-10:30. Gasca, J.M., Moreno-Azanza, M., Ruiz-Omeñaca, J.I., Canudo, J.I. New fossil remains of the ornithopod dinosaur Delapparentia turolensis (Early Cretaceous, Spain) and its phylogenetic position. 10:30-10:45. Bolecsek, C., Wings, O. Europasaurus holgeri the dwarfed dinosaur from the Late Jurassic of Germany. 10:45-11:00. Contreras-Izquierdo, R., Torcida Fernández-Baldor, F., Castilla, M., Rodríguez, L. y Santos, E. Utilidad de la tomografía computerizada (CT) en la preparación paleontológica de un fósil de vertebrado del Cretácico Inferior (Mesozoico) 11:00 Pausa café y Pósters/Cofee break and posters 11:45 Sesión de comunicaciones/oral session 11:45-12:00. Cuenca-Bescós, G., Canudo, J.I., Gasca, J.M., Moreno-Azanza, M Mammals from the time of the dinosaurs, the symmetrodontans from the Early Cretaceous of Spain Programa/Schedule 12:00:12:15. Sellés, A.G. and Vila, B. Laying under stressful conditions: temporal occurrence of pathologic sauropod eggshells in the Late Cretaceous of northeastern Iberian Peninsula 12:15-12:30. Vila, B., Marmi, J., Villalba-Breva, S., Martín-Closas, C. Reconstructing the environment and foraging behaviour of the latest titanosaur herds (Fumanya tracksite, NE Spain) 12:30-12:45. Torcida Fernández-Baldor, F., Canudo, J.I., Contreras, R., Huerta, P., Montero, D., Urién, V. Description of a dorsal vertebra from the sauropod from El Oterillo II (Burgos, Spain) 12:45-13:00. Mocho, P., Royo-Torres, R., Ortega, F., New approach to Lourinhasaurus alenquerensis (Macronaria, Camarasauromorpha) from the Portuguese Upper Jurassic. 13:00 Conferencia Plenaria/Keynote lecture Carole Gee. Sauropod herbivory and the Mesozoic flora. 13:45 Comida/Lunch time 16:00 Conferencia plenaria/keynote lecture Fidel Torcida. Rebbachisauridae: the European exception 16:45 Sesión de comunicaciones. 16:45-17:00. Mocho, P., Ortega, F., Aberasturi, A., Escaso, F. Spinophorosaurus (Sauropoda), a new look inside eusauropod evolution 17:00-17:15. Marmi, J., Villalba-Breva, S., Gomez, B., Martín-Closas, C., Daviero-Gomez, V. Megafossil plant assemblages from the early Maastrichtian of northeastern Iberia challenge the paradigm of the rise to dominance of angiosperms at local scales 17:15-17:30. James, H. K. Palaeontology, Pangaea and Plate Tectonics Data before Models 17:30 Pausa café/cofee break and posters 18:00 Acto de clausura/closing ceremony 18:30 Conferencia Plenaria/Keynote lecture Ronan Allain. A cross-perspective into the Early Cretaceous dinosaur faunas from Europe and Asia: new data on the dinosaurs from the Savannakhet Bassin (Laos) and from the lignitic bone bed of Angeac (France). 19:15 Fin de la sesión 21:30 Cena de clausura/congress dinner. 23:30 Actividades culturales/cultural activities Concierto/Concert Huellas por Jorge Pardo Quartet Horario Sabado día 7 10:00 Excursión a yacimientos de icnitas de la comarca/ Fieldtrip to tracksites in the Salas area 11
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Presentación El Colectivo Arqueológico-Paleontológico de Salas, C.A.S., con la colaboración de diferentes entidades públicas y privadas, organiza desde 1999 reuniones científicas que tratan monográficamente sobre la investigación de los dinosaurios y el medio ambiente en que vivieron. Las Jornadas de Salas tienen como principal objetivo la presentación y discusión de los avances más recientes sobre la Paleontología de dinosaurios, incluyendo lo relacionado con los ecosistemas mesozoicos en los que habitaron, así como la gestión de la conservación y puesta en valor de este patrimonio paleontológico, y su interés en la didáctica de la Ciencia. Esta reunión científica supone el encuentro de investigadores, profesores, estudiantes y otras personas interesadas en la Paleontología del Mesozoico, procedentes de España y otros países. La comarca natural de Salas de los Infantes, rica en fósiles de dinosaurios, impulsa de este modo la función dinamizadora del patrimonio paleontológico de Castilla y León, con el objetivo de que la sociedad de nuestro territorio sea beneficiaria final de los avances del conocimiento científico. Esta convocatoria se produce en un momento difícil para la Ciencia en España, con problemas de financiación de proyectos y grupos de investigación consolidados, y que precisan del apoyo claro de las instituciones. Las Jornadas suponen, en estos momentos con más sentido, una apuesta decidida por la ciencia y el beneficio social que esta supone. Deseamos que todos los asistentes disfruten de la discusión cientifica entre amigos y que estos días sean muy fructiferos tanto para los asistentes como para la paleontología de dinosaurios en general. El Comité organizador. Presentation Since 1999, the Archaeological-Palaeontological Group of Salas (Colectivo Arqueológico-Paleontológico de Salas, C.A.S.), with the collaboration of various public and private institutions, has been organizing scientific meetings focused entirely on dinosaurs, cohabitant animals and the environment in which they lived. The aim of the Salas Conference is the presentation and discussion of the most recent advances in dinosaur palaeontology, including those related to the Mesozoic ecosystems in which they lived, as well as the conservation and publicizing of our palaeontological heritage, and its relevance to the teaching of science. The conference provides a meeting-point for researchers, teachers, students and other people interested in Mesozoic palaeontology, both from Spain and elsewhere. In this way, the Salas de los Infantes area, with its richness of dinosaur fossils, stimulates the power of the palaeontological heritage of Castilla y León to renew and revitalize the region, being the society of the region the ultimate beneficiary of the latest advances in scientific knowledge. The meeting is taking place at a difficult moment for science in Spain, with a weak support from institutions to research projects and groups. At this time, more than ever, the conference represents a decided call for science and for the social benefits that it brings. The organizing comitee wishes all the participants to enjoy the scientific discussion among friends, and that all of these days will be very productive to you, and to the dinosaur palaeontology in general. The organizing comitee. 13
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Conferencias 15
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A cross-perspective into the Early Cretaceous dinosaur faunas from Europe and Asia: new data on the dinosaurs from the Savannakhet Bassin (Laos) and from the lignitic bonebed of Angeac-Charente (France) Allain, R. 1, Rozada. L. 1, Vullo, R. 2 1 Muséum National d Histoire Naturelle, Centre de Recherche sur la Paléobiodiversité et les Paléoenvironnements, UMR 7207 (CNRS/MNHN/UPMC), CP 38, 57 rue Cuvier F-75231 Paris cedex 05, France. rallain@mnhn.fr 2 Université Rennes 1, Géosciences, CNRS UMR 6118, Campus de Beaulieu bat. 15, 263 avenue du Général Leclerc, F-35042 Rennes cedex, France. Abstract Recent fieldworks in the Early Cretaceous from Southeast Asia and Western Europe have yielded new material of continental vertebrate taxa which suggests a complex palaeobiogeographical pattern for the Early Cretaceous and more specifically contacts between Europe and Asia well before the Aptian, the time when iguanodonts seem to diversify in Asia (Russell, 1993; Le Loeuff, 1997). The lignitic deposits from the new Hauterivian-Barremian locality of Angeac-Charente (southwestern France) have yielded a rich (more than 2500 remains have been collected) and diverse vertebrate assemblage (Allain et al., 2011; Neraudeau et al., 2012). Several reptilian taxa, including three turtles, one plesiosaur, four crocodilians and seven dinosaurs (one allosauroid, one ornithomimosaur, one tyrannosauroid, one sauropod, one iguanodont, one hypsilophodont, and one stegosaur) have been identified. Preliminary results indicate that this bone-bed is largely dominated by a new species of ornithomimosaur, represented by at least 20 individuals that have been unearthed from a surface of about 100 square meters. While most of these bones are well preserved, with no abrasion, long ornithomimosaur bones often display post-mortem fractures with sharp edges. Some scratches can be observed on the bone surface. Such observations reveal peculiar taphonomic conditions and are congruent with the hypothesis of a single, short-term burial event possibly followed by trampling. Distinct age-classes are present, with juveniles and sub-adults. This would suggest gregarious habits, a behavior previously reported for this group of herbivorous theropod dinosaurs. The unambiguous fossil record of European ornithomimosaurs has been so far restricted to Pelecanimimus from the Spanish Barremian locality of Las Hoyas, a primitive toothed form known only by a single specimen. Comparisons with material from the Wealden area (e.g., Thecocoelurus and Valdoraptor) suggest that ornithomimosaurs were also present in the Early Cretaceous of England. There are now many elements in common between European and Asian assemblages in the mid Early Cretaceous, including several dinosaur groups such as the Baryonychinae (Asia: Buffetaut et al., 2005; Allain et al., 2012; Europe: Charig & Milner, 1997), the Ornithomimosauria (see above), various basal Titanosauriformes and Carcharodontosauria, but also trigonioidoid non-marine bivalves (Sha, 2010; Delvene et al., 2011), goniopholidid and atoposaurid crocodyliforms (Lauprasert et al., 2007; 2011), istiodactylid pterosaurs (Andres & Ji, 2006) and gobiconodontid mammals (Cuenca-Bescós & Canudo, 2003). These rather uniform assemblages may be linked to the very low sea-levels of the earliest Cretaceous which would have allowed exchanges between Euramerica and Asia. Keywords: Dinosaurs, Theropoda, Sauropoda, Iguanodontia, Early Cretaceous, Europe, Asia. References Allain, R., Vullo R., Leprince, A., Neraudeau D., Tournepiche, J.-F. (2011): An ornithomimosaur-dominated bonebed from the Early Cretaceous of Southwestern France. Journal of Vertebrate Paleontology. (31, supp. 6), 61. Allain, R., Xaisanavong, T., Richir, P., Khentavong, B. (2012): The first definitive Asian spinosaurid (Dinosauria: Theropoda) from the Early Cretaceous of Laos. Naturwissenschaften. (99). 369-377. 17
Andres, B., Ji, Q. (2006): A new species of Istiodactylus (Pterosauria, Pterodactyloidea) from the Lower Cretaceous of Liaoning, China. Journal of Vertebrate Paleontology. (26), 70-78. Buffetaut, E., Suteethorn, V., Le Loeuff, J., Khansubha, S., Tong, H., Wongko, K. (2005): The Dinosaur fauna from the Khok Kruat Formation (Early Cretaceous) of Thailand. Proceedings of the International Conference on Geology, Geotechnology and Mineral Resources of Indochina (GEOINDO), Khon Kaen, Thailand, 575-581. Charig, A.J., Milner, A.C. (1997): Baryonyx walkeri, a fish-eating dinosaur from the Wealden of Surrey. Bulletin of the Natural History Museum London. (53), 11-70. Cuenca-Bescós, G., Canudo, J.I. (2003): A new gobiconodontid mammal from the Early Cretaceous of Spain and its palaeogeographic implications. Acta Paleontologica Polonica. (48), 575-582. Delvene, G., Munt, M., Sender, L.M. (2011): Iberanaia iberica: The first record of the Trigonioidoidea Bivalvia: Unionoida from the Lower Cretaceous of Teruel, Spain. Cretaceous Research. (32), 591-596. Lauprasert, K., Cuny, G., Buffetaut, E., Thirakhupt, K., Suteethorn, V. (2007): Siamosuchus phuphokensis, a new goniopholidid from the Early Cretaceous (ante-aptian) of northeastern Thailand. Bulletin de la Société Géologique de France. (178), 201-216. Lauprasert, K., Laojumpon, C., Saenphala, W., Cuny, G., Thirakhupt, K., Suteethorn, V. (2011): Atoposaurid crocodyliforms from the Khorat Group of Thailand: first record of Theriosuchus from Southeast Asia. Paläontologische Zeitschrift. (85), 37-47. Le Loeuff, J. (1997): Biogeography. In: Currie, P.J., Padian, K. (eds.). Encyclopedia of Dinosaurs, Academic Press, 51-56. Néraudeau, D., Allain, R., Ballèvre, M., Batten, D.J., Buffetaut, E., Colin, J.P., Dabard, M.P., Daviero-Gomez, V., El Albani, A., Gomez, B., Grosheny, D., Le Lœuff, J., Leprince, A., Martin-Closas, C., Masure, E., Mazin, J.-M., Philippe, M., Pouech, J., Tong, H., Tournepiche, J.F., Vullo, R. (2012): The Hauterivian Barremian lignitic bone bed of Angeac (Charente, SW France): stratigraphical, palaeobiological and palaeogeographical implications. Cretaceous Research. (37), 1-14 Russell, D.A. (1993): The role of Central Asia in dinosaurian biogeography. Canadian Journal of Earth Sciences. (30), 2002-2012. Sha, J. (2010): Historical distribution patterns of trigonioidids (non-marine Cretaceous bivalves) in Asia and their palaeogeographic significance. Proceedings of the Royal Society B. (277), 277-283. 18
Ten years digging up dinosaurs in Patagonia: the Ibero-Patagonian connection in the Mesozoic Canudo, J. I. 1 Salgado, L. 2 Carballido, J. 3 Garrido, A. 4 1: Grupo Aragosaurus-IUCA (Instituto Universitario de Ciencias Ambientales de Aragón), http://www.aragosaurus.com, Paleontología. Facultad de Ciencias. C/Pedro Cerbuna 12. 50009 Zaragoza (Spain). jicanudo@unizar.es 2: CONICET, Universidad de Río Negro, C/ Isidro Lobo y Belgrano. 8332 General Roca, Río Negro, Argentina. 3: CONICET, Museo Paleontológico Egidio Feruglio, Fontana 140, 9100 Trelew, Argentina. 4. Museo Provincial de Ciencias Naturales Prof. Dr. Juan A. Olsacher, Etcheluz y Ejército Argentino, 8340 Zapala, Neuquén, Argentina Abstract The Salas congress serves as a meeting point where researchers from different parts of the world can devise projects in common. A good example of this is the series of joint research projects that we have been undertaking for more than ten years now. It started when two of us coincided at the 2001 Salas congress, where we raised a number of palaeobiogeographial questions that called for research into the vertebrates of Patagonia and Iberia (Canudo and Salgado, 2003). Researchers from the University of Zaragoza and various Argentinean institutions (Museo Olsacher de Zapala, Museo de Rincón de los Sauces, Museo Egidio Feruglio de Trelew, Universidad de Comahue, Universidad of Río Negro, Universidad de La Plata, Conicet) have taken part in these research projects. They have been pursued for more than ten years, financed mainly by the National Research Plan of the Government of Spain, but also supported by private enterprises such as Repsol and Petrobras, museums such as the Museo of Rincón de los Sauces and of Zapala, and institutions such as the Mining Office of Neuquén and Endemás of Cipolletti. The fundamental idea was to use terrestrial vertebrates especially dinosaurs to infer the palaeobiogeographical relationships that existed between Iberia and Patagonia during the Cretaceous. South America and Africa were joined for most of the Lower Cretaceous, and there were also connections with Europe (Canudo et al., 2009). Standard models locate the final separation of Africa and South America at the end of the Lower Cretaceous or at the beginning of the Upper Cretaceous, with the complete opening of the South Atlantic. One of the objectives of the project is precisely to pinpoint this definitive isolation. The presence in the Campanian- Maastrichtian of Europe of derived taxa of titanosaurian sauropods that are close to Argentinean taxa makes the scenario more complicated, since South America had apparently been isolated since the beginning of the Upper Cretaceous. In this communication we shall focus only on the work carried out in the region of Patagonia in Argentina. The phylogenetic proximity of taxa that are now separated by geographical barriers (such as oceans separating distinct continents) is one of the criteria that are commonly used as proof of geographical connections in the past. If we find the same species on both sides of the barrier constituted by the ocean, the conclusion is immediate: when these two organisms were alive, the barrier did not exist. However, this is an ideal situation; in many cases it is representatives of the same supra-specific group that are found, but these may have undergone vicariant evolution within the two landmasses. Clarifying just such questions is one of the main lines of the project: distinguishing endemic taxa (both in Europe and in South America) from those taxa that exist in a close phylogenetic relation and that may have played a role in the dispersals from Gondwana to Laurasia or vice versa. From the outset this project has focused on a search for new specimens, above all in critical periods such as the upper part of the Lower Cretaceous and the lower part of the Upper Cretaceous. Prospection and excavation field campaigns (generally one a year) have been undertaken in the provinces of Neuquén and Río Negro. The field projects have generally been carried out in new areas and with little prior information. The 19
first projects were undertaken in the Upper Cretaceous of the areas of El Anfiteatro (Río Negro) and Rincón de los Sauces (Neuquén). In El Anfiteatro a great quantity of fragmentary vertebrate material was found, shedding light on the palaeobiodiversity of the area (Salgado et al., 2009). In Rincón de los Sauces the remains of various titanosaurian sauropods have been excavated, one of which was described as Petrobrasaurus puestohernandezi from the Coniacian-Santonian of the Plottier Formation (Filippi et al., 2011). This period is particularly interesting as this was when the definitive separation of Africa and South America took place. Petrobrasaurus is a form that is related to other Argentinean titanosaurs and unrelated to the European ones. A group of special interest to the palaeobiogeographical debate is the rebbachisaurids. These are basal diplodocimorphs represented in South America, Africa and Iberian Peninsula. A particularly interesting area is the west of the province of Neuquén with large outcrops from the Lower Cretaceous Upper Cretaceous boundary. A revision has been undertaken of the type area and material of Rayososaurus agrioensis from the Candeleros Formation (lower Cenomanian), initially considered to date to the Lower Cretaceous (Carballido et al., 2010). Furthermore, the new taxon Comahuesaurus windhauseni has been described on the basis of postcranial material from the Aptian-Albian of the Lohan Cura Formation (Carballido et al., 2012). These rebbachisaurids seem to form part of an exclusively South American group. In the course of our field work, we have for the first time discovered sauropod remains in the Rayoso Formation. These belong to a semi-articulated individual, including cranial elements, and two juvenile individuals (Salgado et al., 2012). This material is currently under preparation, but has been initially assigned to cf. Zapalasaurus. This taxon is of palaeobiogeographical interest because in phylogenetic proposals it is grouped in the same clade as Nigersaurus (Africa) and Demandasaurus (Spain). Keywords: Dinosaurs, Paleobiogeography, Cretaceous, Iberian Peninsula. Argentina References Canudo J.I., Barco J.L., Pereda-Suberbiola X., Ruiz-Omeñaca J.I., Salgado, L., Torcida Fernández-Baldor F., Gasulla J.M. (2009): What Iberian dinosaurs reveal about the bridge said to exist between Gondwana and Laurasia in the Early Cretaceous. Bulletin de la Sociéte Geológique de France. (180), 5-11. Canudo, J.I., Salgado, L. (2003): Los dinosaurios del Neocomiense (Cretácico inferior) de la Península Ibérica y Gondwana occidental: implicaciones paleobiogeográficas. En: Dinosaurios y otros reptiles mesozoicos de España. Instituto de Estudios Riojanos. (26), 251-268. Carballido, J.L., Garrido, A.C., Canudo, J.I., Salgado, L. (2010): Redescription of Rayososaurus agrioensis Bonaparte (Dinosauria, Diplodocoidea), a rebbachisaurid of the early Upper Cretaceous of Neuquén. Geobios. (43), 493-502. Carballido, J.L., Salgado, L., Pol, D., Canudo, J.I., Garrido, A. (2012): A new basal rebbachisaurid (Sauropoda, Diplodocoidea) from the Early Cretaceous of the Neuquén Group; evolution and biogeography of the group. Historical Biology. (24), 631-654. Filippi L., Canudo J.I., Salgado L., Garrido A., García R., Cerda I., Otero, A. (2011): A new sauropod from the Plottier Formation of Patagonia (Argentina). Geologica Acta. (9), 1-12. Salgado, L., Canudo, J.I., Garrido, A.C., Carballido, J.L. (2012): Evidence of gregariousness in Rebbachisauridae (Dinosauria, Sauropoda, Diplodocoidea) in the Early Cretaceous of Neuquén (Rayoso Formation), Patagonia, Argentina. Journal of Vertebrate Paleontology. (32), 603-613. Salgado L., Canudo J.I., Garrido A.C., Ruiz-Omeñaca J.I., García R.A., de la Fuente M.S., Barco J.L., Bollati R. (2009): Upper Cretaceous vertebrates from El Anfiteatro (Río Negro, Patagonia, Argentina). Cretaceous Research. (30), 767-784. 20